​Education
1978-1984 ; Faculty of Medicine, Kagoshima University
1984-1988 ; Graduate course, Department of Biochemistry,
Faculty of Medicine, Kagoshima University,
Awarded the degree of PhD in Biochemistry.

Research and professional experience
1987-1989; Special Researcher of Japan Society for the Promotion of Science, Tokushima University
1989-1991; Post-doctoral fellow, Department of Biochemistry,
University of Utah
1991-1993; Research Associate, Howard Hughes Medical Institute,
University of Utah
1993-1997; Research Associate, Department of Human Genetics,
University of Utah
1997-2000; Associate Professor, Institute for Molecular and Cellular Regulation, Gunma University
2000-2006; Associate Professor, Institute of Medical Science, University of Tokyo
2006-2010; Professor, School of Medicine, University of Tokai
2008-2010; Director, Institute of Medical Sciences, University of Tokai
2010-present; Professor, National Institute of Genetics